C3 and C4 pathways:
It was one time thought that all green plants fix CO2 by Calvin cycle only. Here, we make out that specific plants fix CO2 in a dissimilar photosynthetic mechanism termed as C4 pathway. Hatch and Slack discovered that 4C compounds like malate, oxaloaceticacid, and aspartate were the first created compounds, while the leaves of sugarcane were displayed to 14CO2 for one second. Thus, sugarcane is an instance for C4 plant. While the leaves of rice plant are diaplayed to 14CO2, 3C compound called phosphoglyceric acid is created. Thus, rice plant is an instance for C3 plant.
In C3 plants, photosynthesis takes place just only in mesophyll cells. We previously learnt that photosynthesis has two sorts of reactions –
1. Light reactions and 2. Dark reactions (Calvin cycle).
In the first type of reaction that is light reactions ATP and NADPH2 are generated and oxygen is released like a byproduct. CO2 is reduced to carbohydrates through dark reactions. In C3 plants both the first reaction that is light reactions and second reaction dark reactions take place in mesophyll cells, while in C4 plants, the technique of photosynthesis needs two sorts of photosynthetic cells –
1. mesophyll cells and 2. bundle sheath cells.
The C4 plants consist of dimorphic chloroplast that is chloroplasts in mesophyll cells are granal (with grana) while in bundle sheath chloroplasts are agranal (with no grana). The existence of two sorts of cells leads to segregation of photosynthetic work that is light reactions and dark reactions separately.
Diagram: Hatch-Slack pathway of CO2 fixation
(PEP = Phosphoenol pyruvic acid, OAA = Oxalo acetic acid)
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