Two genes control color in corn snakes as follows: O_B_ snakes are brown, O_bb are orange, ooB_ are black, and oobb are albino. An orange snake was mated to a black snake, and a large number of F1 progeny were obtained, all of which were brown. When the F1 snakes were mated to one another, they produced 100 brown offspring, 25 orange, 22 black and 13 albino. What are the genotypes of the F1 snakes?