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Effects of ozone exposure to aspen and birch

During the past 12 years, scientists have been exposing stands of aspen and birch to elevated levels of atmospheric O? at a facility (Aspen FACE) near Rhinelander, WI. The relative effects of O? exposure on stand growth has varied considerably among years. Based on the data below, what is a plausible explanation for differences in O? effects observed in Year 4 versus Year 5?

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