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How female is approachable to copulation

Noramlly how does a male animal recognize that the female is approachable to copulation?

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In the majority of vertebrate species with interior fecundation the females have reproductive cycles with fertile stages. For this period the female secretes pheromones (odoriferous substances which attract male of the species) from mucosae and skin. The existence of the male individual and its pheromones too stimulates the release of pheromones by female. (Several animals also use pheromones for territorial demarcation in addition for signal transmission among individuals concerning the location of dangers and food.)

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